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Hero Souls are items that provide global DPS bonuses and can be used as currency to summon and level up Artifacts.
Each hero soul provides a +10% bonus to the Damage Per Second.
The hero Brath the Merciless has a skill called DARK RITUAL. This resets the game and starts over, with a world just like the current one, but granting a number of Hero Souls depending on the following:
For every 200 hero levels across all heroes, you get a single Hero Soul for your next ascension. In the first few ascensions, this will likely be the bulk of the Hero Soul earnings.
Starting at level 100, all boss levels have a chance of spawning a Specter the first time you enter that level on each ascension. The Specter awards a number of hero souls for your next Dark Ritual depending on the level they are defeated on. Remember, the souls are only added to the number of souls you get from resetting. It doesn't add to your current amount of souls immediately. Specter can only be encountered once per floor each ascension, then will be replaced by the normal versions.
The base encounter rate for Specters is 25%.